When the intine had been formed, it revealed remarkable ATPase activty, and the pollens were full of mitochondria with developed cristae where cytochrome oxidase activity was observed.

• 而不育系花药，随着花粉的发育，花粉粒质膜上的ATP酶活性增加不明显，且内壁发育异常，在内壁中几乎没有ATP酶活性出现。