Using radioactively labeled elements he demonstrated that the region of maximum absorption of water is a few centimeters behind the root tip and that quantities of minerals are absorbed passively in the transpiration stream.

• 利用放射性标记元素证明了水分吸收的主要部位在根尖以上几厘米，矿物质随蒸腾作用被动的吸收入植物体内。