They are the dominant organisms in aeratedsoils (Frey et al.1999), typically accounting for 10%?60% of the biomass inforest litter (Newell 1992; Metting 1993).

• 在含有氧合的土壤中它们也是优势生物，典型的证明是在森林的枯枝落叶层中有生物量的10%25~60%25。