These glycosaminoglycan (GAG) chains are long, linear carbohydrate polymers that are negatively charged under physiological conditions, due to the occurrence of sulphate and uronic acid groups.

• 这些粘多糖（GAG）链是长条线状的碳水化合物聚合体，在正常生环境下，由于硫酸根和糖醛酸根的存在而处于负电位。