The process of spore germination in D. varia is attributed to Vittaria-type, and that of gametophyte development belongs to Aspidium-type.This feature is consistent with that in Dryopteris Adanson.

• 孢子萌发为书带蕨型，原叶体发育为三叉蕨型，符合鳞毛蕨属配子体发育的特征。