The number and shape of pinnatifid-lobes of leaflets, morphology of involucral bracts, and color of flowers are shown to be the most useful characters for separating Taiwanese species.

• 本研究显示，小叶的羽状裂瓣数目的形状、緫苞片的形态、以及花的颜色，提供台湾产种类相当重要之分类依据。