The anther wall layers consist of the epidermis, the endothecium of one cell-layer, the middle layers of two cells and the tapetum of one-layer.

• 花药壁包含，一层细胞的表皮层、一层细胞的内壁层、两层细胞的中间层及一层细胞的营养层，此药壁发育方式可归类为基本型。