Mueller and his colleagues tested tattooing by vaccinating mice with a protein fragment of human papillomavirus, or HPV, a sexually transmitted virus that causes cervical cancer.

• 缪勒和和他的同事们，通过接种了一种人类乳头瘤病毒或一种能够引起子宫颈癌的性传播病毒HPV蛋白片段疫苗的老鼠做了实验。