Less well known, but perhaps of equal evolutionary significance, are the studies on early insect flight, on stem tetrapods from the Upper Devonian, and the early stages of whale evolution.

• 对早期昆虫的飞行、晚泥盆世羽轴类四脚动物和鲸进化的早期阶段的研究不太为人所知，但也许有着同样重要的进化意义。