In chloroplasts thylakoid membranes became loose gradually, whereas the number and size of osmiophilic granule increased till the chloroplasts disappeared.

• 叶片衰老过程中叶绿体类囊体膜片层逐渐松散，直至膜结构逐渐解体，叶绿体内油脂颗粒增大增多，最终解体。