Besides, the epidermis characters showed that Cryptogramma, Onychium and Acrostichum are unique.And more study of the three genera is needed to understand the relationships among these genera.

• 此外，珠蕨属、金粉蕨属和卤蕨属的叶表皮特徵和其他属均不相似，而它们在凤尾蕨科内的分类地位值得进一步研究。