According to the properties of litter substrate, the enzymes are classified into cellulolytic enzymes, ligninolytic enzymes, proteolytic enzymes and phosphatase.

• 根据森林凋落物底物性质的不同，将凋落物分解酶分为纤维素分解酶类、木质素分解酶类、蛋白水解酶类和磷酸酶类。